BURNS, Ore. (AP) — Surrounded by FBI agents in armored vehicles, the last four occupiers of a national nature preserve surrendered Thursday, and a leader in their movement who organized a 2014 standoff with authorities was criminally charged in federal court.

The holdouts were the last remnants of a larger group that seized the Malheur National Wildlife Refuge nearly six weeks ago, demanding that the government turn over the land to locals and release two ranchers imprisoned for setting fires. For the first time since Jan. 2, the federal land was fully under the control of the U.S. government.

Meanwhile, Cliven Bundy, who was at the center of the 2014 standoff at his ranch in Nevada, was arrested late Wednesday in Portland after encouraging the Oregon occupiers not to give up. Bundy is the father of Ammon Bundy, the jailed leader of the Oregon occupation.